So, his crown doesn't match any recognized crown type. Which is very interesting, but considering we don't even know what exact genes are involved in recognized crown types it would be hard to determine what genes are involved in an abnormal comb. I think it's worth mentioning that gene...
Comb: there is a lot we don't know about comb genetics, but here's what I do know.
There are 5 known locusts, containing 11 alleles, that we know of, and they control comb type:
1. Pea Comb Locus, P, p+
2. Rose Comb Locus, R r+
3. Duplex Comb Locus, Dv, Dc, d+
4. Breda Comb Locus, bd, Bd+
5...